Not only do hypostomes rarely appear in fossil shows, they are not widely
recognized. Often or in may cases totally lost in molting and death,
hypostotmes were part of the feeding apparatus of these primitive arthropods,
were lodged under the cephalon.
Given the great variety of trilobites over the hundreds of millions of years
of their existence, diets would have varied a great deal. Forms and textures
would be subject to change according to orders, families, and even genera.
This limited selection, spanning time from an earliest Ordovician olenid
to a middle Devonian lichid, gives a sampling of the notable differences
that make for intriguing study. It is hard to believe that the illustrations
of the
Hoekaspis and the
Asaphellus
are both from asaphids. The fingerprint-like ridge marks
on the Hoekaspis may have facilitated food ingestion. The
main body of the cephalon was eroded disclosing this setting for its
under-anatomy.
As often as not, hypostomes are found displaced from other parts
of their anatomy. The large specimen of a
Fenestraspis amauta
was preserved through
the ages in a concretion all to itself. The bulbous main body would have
been on the underside of the glabella... Not surprisingly no head that size
has been collected.